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Carbohydrates overview

Carbohydrates are chemical compoundstogether by a covalent glycosidic bond.
that contain oxygen, hydrogen, andThe binding between the two sugars
carbon atoms, and no other elements.results in the loss of a hydrogen atom
They consist of monosaccharide sugars of(H) from one molecule and a hydroxyl
varying chain lengths and that have thegroup (OH) from the other.
general chemical formula Cn(H2O)n or areThe most common disaccharides are
derivatives of such. The smallest valuesucrose (cane or beet sugar - made from
for "n" is 3. A 3-carbon sugar isone glucose and one fructose), lactose
referred to as a triose, whereas a(milk sugar - made from one glucose and
6-carbon sugar is called a hexose (seeone galactose) and maltose (made of two
monosaccharides below).glucoses). The formula of these
Certain carbohydrates are an importantdisaccharides is C12H22O11.
storage and transport form of energy inOligosaccharides and polysaccharides
most organisms, including plants andOligosaccharides and polysaccharides are
animals. Carbohydrates are classified bycomposed of longer chains of
their number of sugar units:monosaccharide units bound together by
monosaccharides (such as glucose andglycosidic bonds. The distinction
fructose), disaccharides (such asbetween the two is based upon the number
sucrose and lactose), oligosaccharides,of monosaccharide units present in the
and polysaccharides (such as starch,chain. Oligosaccharides typically
glycogen, and cellulose).contain between three and nine
Pure carbohydrates contain carbon,monosaccharide units, and
hydrogen, and oxygen atoms, in a 1:2:1polysaccharides contain greater than ten
molar ratio, giving the general formulamonosaccharide units.
Cn(H2O)n. (This applies only toDefinitions of how large a carbohydrate
monosaccharides, see below, although allmust be to fall into each category vary
carbohydrates have the more generalhowever.
formula Cn(H2O)m.) However, manyOligosaccharides are found as a common
important "carbohydrates" deviate fromform of protein posttranslational
this, such as deoxyribose and glycerol,modification. Polysaccharides represent
although they are not, in the strictan important class of biological
sense, carbohydrates. Sometimespolymer.
compounds containing other elements areExamples include starch, cellulose,
also counted as carbohydrates (e.g.chitin and glycogen.
chitin, which contains nitrogen).Nutrition Strictly speaking,
The simplest carbohydrates arecarbohydrates are not necessary for
monosaccharides, which are smallhuman nutrition because proteins can be
straight-chain aldehydes and ketonesconverted to carbohydrates. The
with many hydroxyl groups added, usuallytraditional diet of some cultures
one on each carbon except the functionalconsists of very little carbohydrate,
group. Other carbohydrates are composedand these people remain relatively
of monosaccharide units and break downhealthy.[citation needed] However,
under hydrolysis. These may becarbohydrates require less water to
classified as disaccharides,digest than proteins or fats and are the
oligosaccharides, or polysaccharides,most common source of energy.
depending on whether they have two,Proteins and fat are vital building
several, or many monosaccharide units.components for body tissue and cells,
Monosaccharides Monosaccharides may beand thus it could be considered
divided into aldoses, which have anadvisable not to deplete such resources
aldehyde group on the first carbon atom,by necessitating their use in energy
and ketoses, which typically have aproduction. Carbohydrates, like
ketone group on the second. They mayproteins, contain 4 kilocalories per
also be divided into trioses, tetroses,gram while fats contain 9 kilocalories
pentoses, hexoses, and so forth,and alcohol contains 7 kilocalories per
depending on how many carbon atoms theygram.
contain. For instance, glucose is anBased on evidence for risk of heart
aldohexose, fructose a ketohexose, anddisease and obesity, the Institute of
ribose an aldopentose.Medicine recommends that American and
Further, each carbon atom that supportsCanadian adults get between 40-65% of
a hydroxyl group (except for the firstdietary energy from carbohydrates.[1]
and last) is optically active, allowingThe Food and Agriculture Organization
a number of different carbohydrates withand World Health Organization jointly
the same basic structure. For instance,recommend that national dietary
galactose is an aldohexose but hasguidelines set a goal of 55-75% of total
different properties from glucoseenergy from carbohydrates.[2] Foods high
because the atoms are arrangedin carbohydrates Breads, pastas,
differently.potatoes, bran and cereals are all high
The straight-chain structure describedin carbohydrates.
here is only one of the forms aClassification Dietitians and
monosaccharide may take. The aldehyde ornutritionists commonly classify
ketone group may react with a hydroxylcarbohydrates as simple (monosaccharides
group on a different carbon atom to formand disaccharides) or complex
a hemiacetal or hemiketal, in which case(oligosaccharides and polysaccharides),
there is an oxygen bridge between thedepending on their chemical structure.
two carbon atoms, forming a heterocyclicThe term complex carbohydrate was first
ring. Rings with five and six atoms areused in the Senate Select Committee
called furanose and pyranose forms andpublication Dietary Goals for the United
exist in equilibrium with theStates (1977), where it denoted "fruit,
straight-chain form.vegetables and whole-grains". Dietary
It should be noted that the ring formguidelines generally recommend that
has one more optically active carboncomplex carbohydrates and nutrient-rich
than the straight-chain form, and so hassimple carbohydrates such as fruit and
both an alpha and a beta form, whichdairy products should make up the bulk
interconvert in equilibrium. However,of carbohydrate consumption. The USDA's
the carbohydrate may further react withDietary Guidelines for Americans 2005
an alcohol to form an acetal or ketal,dispenses with the simple/complex
in which case the two forms becomedistinction, instead recommending
distinct.fiber-rich foods and whole grains.
This is the basic type of link betweenThe glycemic index and glycemic load
the monosaccharide units of largersystems are popular alternative
carbohydrates.classification methods which rank
Disaccharides Disaccharides are composedcarbohydrates based on their effect on
of two monosaccharide units boundblood glucose levels.



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