Carbohydrates overview

Carbohydrates are chemical compounds that containglycosidic bond. The binding between the two sugars
oxygen, hydrogen, and carbon atoms, and no otherresults in the loss of a hydrogen atom (H) from one
elements. They consist of monosaccharide sugars ofmolecule and a hydroxyl group (OH) from the other.
varying chain lengths and that have the generalThe most common disaccharides are sucrose (cane
chemical formula Cn(H2O)n or are derivatives of such.or beet sugar - made from one glucose and one
The smallest value for "n" is 3. A 3-carbon sugar isfructose), lactose (milk sugar - made from one glucose
referred to as a triose, whereas a 6-carbon sugar isand one galactose) and maltose (made of two
called a hexose (see monosaccharides below).glucoses). The formula of these disaccharides is
Certain carbohydrates are an important storage andC12H22O11.
transport form of energy in most organisms, includingOligosaccharides and polysaccharides
plants and animals. Carbohydrates are classified byOligosaccharides and polysaccharides are composed
their number of sugar units: monosaccharides (such asof longer chains of monosaccharide units bound
glucose and fructose), disaccharides (such as sucrosetogether by glycosidic bonds. The distinction between
and lactose), oligosaccharides, and polysaccharidesthe two is based upon the number of monosaccharide
(such as starch, glycogen, and cellulose).units present in the chain. Oligosaccharides typically
Pure carbohydrates contain carbon, hydrogen, andcontain between three and nine monosaccharide units,
oxygen atoms, in a 1:2:1 molar ratio, giving the generaland polysaccharides contain greater than ten
formula Cn(H2O)n. (This applies only tomonosaccharide units.
monosaccharides, see below, although allDefinitions of how large a carbohydrate must be to fall
carbohydrates have the more general formulainto each category vary however.
Cn(H2O)m.) However, many important "carbohydrates"Oligosaccharides are found as a common form of
deviate from this, such as deoxyribose and glycerol,protein posttranslational modification. Polysaccharides
although they are not, in the strict sense,represent an important class of biological polymer.
carbohydrates. Sometimes compounds containingExamples include starch, cellulose, chitin and glycogen.
other elements are also counted as carbohydratesNutrition Strictly speaking, carbohydrates are not
(e.g. chitin, which contains nitrogen).necessary for human nutrition because proteins can
The simplest carbohydrates are monosaccharides,be converted to carbohydrates. The traditional diet of
which are small straight-chain aldehydes and ketonessome cultures consists of very little carbohydrate, and
with many hydroxyl groups added, usually one on eachthese people remain relatively healthy.[citation needed]
carbon except the functional group. OtherHowever, carbohydrates require less water to digest
carbohydrates are composed of monosaccharidethan proteins or fats and are the most common
units and break down under hydrolysis. These may besource of energy.
classified as disaccharides, oligosaccharides, orProteins and fat are vital building components for body
polysaccharides, depending on whether they have two,tissue and cells, and thus it could be considered
several, or many monosaccharide units.advisable not to deplete such resources by
Monosaccharides Monosaccharides may be dividednecessitating their use in energy production.
into aldoses, which have an aldehyde group on the firstCarbohydrates, like proteins, contain 4 kilocalories per
carbon atom, and ketoses, which typically have agram while fats contain 9 kilocalories and alcohol
ketone group on the second. They may also becontains 7 kilocalories per gram.
divided into trioses, tetroses, pentoses, hexoses, andBased on evidence for risk of heart disease and
so forth, depending on how many carbon atoms theyobesity, the Institute of Medicine recommends that
contain. For instance, glucose is an aldohexose,American and Canadian adults get between 40-65%
fructose a ketohexose, and ribose an aldopentose.of dietary energy from carbohydrates.[1] The Food
Further, each carbon atom that supports a hydroxyland Agriculture Organization and World Health
group (except for the first and last) is optically active,Organization jointly recommend that national dietary
allowing a number of different carbohydrates with theguidelines set a goal of 55-75% of total energy from
same basic structure. For instance, galactose is ancarbohydrates.[2] Foods high in carbohydrates Breads,
aldohexose but has different properties from glucosepastas, potatoes, bran and cereals are all high in
because the atoms are arranged differently.carbohydrates.
The straight-chain structure described here is only oneClassification Dietitians and nutritionists commonly
of the forms a monosaccharide may take. Theclassify carbohydrates as simple (monosaccharides
aldehyde or ketone group may react with a hydroxyland disaccharides) or complex (oligosaccharides and
group on a different carbon atom to form apolysaccharides), depending on their chemical structure.
hemiacetal or hemiketal, in which case there is anThe term complex carbohydrate was first used in the
oxygen bridge between the two carbon atoms,Senate Select Committee publication Dietary Goals for
forming a heterocyclic ring. Rings with five and sixthe United States (1977), where it denoted "fruit,
atoms are called furanose and pyranose forms andvegetables and whole-grains". Dietary guidelines
exist in equilibrium with the straight-chain form.generally recommend that complex carbohydrates
It should be noted that the ring form has one moreand nutrient-rich simple carbohydrates such as fruit and
optically active carbon than the straight-chain form, anddairy products should make up the bulk of
so has both an alpha and a beta form, whichcarbohydrate consumption. The USDA's Dietary
interconvert in equilibrium. However, the carbohydrateGuidelines for Americans 2005 dispenses with the
may further react with an alcohol to form an acetal orsimple/complex distinction, instead recommending
ketal, in which case the two forms become distinct.fiber-rich foods and whole grains.
This is the basic type of link between theThe glycemic index and glycemic load systems are
monosaccharide units of larger carbohydrates.popular alternative classification methods which rank
Disaccharides Disaccharides are composed of twocarbohydrates based on their effect on blood glucose
monosaccharide units bound together by a covalentlevels.